[Life of Pi - Rotten Tomatoes](^2^)

Assuming the true distribution is exponential, this would suggest that contraction is triggered by spontaneous fluctuations that occur at a constant rate in time. From observation of movies of filament arrangements, a likely candidate is the transient void formation frequently observed near the outer wall, where nearby filaments are attached purely by motors at their [+]-ends and not along their length. Such voids, when large enough, lead to a 'hinge'-like mechanism in which the void expands and one section of the polarity field inverts, leading to the semi-aster.

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Movie showing the transition from the initial conditions to a spindle steady state. System parameters are identical to Figure 2(a). The colour scheme for this and all movies are as follows: Filaments are light (dark) near their plus (minus) ends; the blue-green hue is to aid visualisation and has no significance. Motors are coloured in red. 2ff7e9595c